Photos by Orest Dorosh – Anthrax’s lineup has changed numerous times over their career, leaving Ian as the only constant member of the band. The band has had a revolving cast of singers and lead guitarists, including Neil Turbin, John Bush, Dan Spitz and Rob Caggiano. Since 2013, the band has consisted of Ian, drummer Charlie Benante, bassist Frank Bello, vocalist Joey Belladonna and lead guitarist Jonathan Donais. Ian and Benante (who replaced one-time drummer Greg D’Angelo in 1983) are the only two members to appear on every album, while Bello has been a member of Anthrax since 1984, replacing original bassist Dan Lilker. After two stints in Anthrax, from 1984 to 1992 and again from 2005 to 2007, Belladonna returned to the band in 2010.

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